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@article{Jinam2015UniqueCO,
  title={Unique characteristics of the Ainu population in Northern Japan},
  author={Timothy A. Jinam and Hideaki Kanzawa-Kiriyama and Ituro Inoue and Katsushi Tokunaga and Keiichi Omoto and Naruya Saitou},
  journal={Journal of Human Genetics},
  year={2015},
  volume={60},
  pages={565-571},
  url={https://api.semanticscholar.org/CorpusID:205166287}
}

It is found that the Ainu are genetically different from Mainland Japanese living in Tohoku, the northern part of Honshu Island, and estimated haplotypes of some Ainu individuals suggested relatively long-term admixture with MainlandJapanese.

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