Giant kelp, Macrocystis pyrifera, increases faunal diversity through physical engineering - PubMed
- ️Mon Jan 01 2018
Giant kelp, Macrocystis pyrifera, increases faunal diversity through physical engineering
Robert J Miller et al. Proc Biol Sci. 2018.
Abstract
Foundation species define the ecosystems they live in, but ecologists have often characterized dominant plants as foundational without supporting evidence. Giant kelp has long been considered a marine foundation species due to its complex structure and high productivity; however, there is little quantitative evidence to evaluate this. Here, we apply structural equation modelling to a 15-year time series of reef community data to evaluate how giant kelp affects the reef community. Although species richness was positively associated with giant kelp biomass, most direct paths did not involve giant kelp. Instead, the foundational qualities of giant kelp were driven mostly by indirect effects attributed to its dominant physical structure and associated engineering influence on the ecosystem, rather than by its use as food by invertebrates and fishes. Giant kelp structure has indirect effects because it shades out understorey algae that compete with sessile invertebrates. When released from competition, sessile species in turn increase the diversity of mobile predators. Sea urchin grazing effects could have been misinterpreted as kelp effects, because sea urchins can overgraze giant kelp, understorey algae and sessile invertebrates alike. Our results confirm the high diversity and biomass associated with kelp forests, but highlight how species interactions and habitat attributes can be misconstrued as direct consequences of a foundation species like giant kelp.
Keywords: competition; ecosystem engineering; facilitation; foundation species; kelp forest; subtidal.
© 2018 The Author(s).
Conflict of interest statement
We have no competing interests.
Figures

Piecewise SEM model of the effect of giant kelp biomass and sea urchin biomass on biomass and diversity order 1 (see Material and methods) of four taxonomic functional groups, represented by the blue boxes: (1) understorey macroalgae, (2) sessile invertebrates, (3) mobile grazers and (4) mobile predators. Effects of substrate on the biomass of sessile groups are also shown. Arrows represent unidirectional relationships among variables. Blue arrows denote positive relationships, and orange arrows negative relationships. Arrows for non-significant paths (p ≥ 0.05) are not shown. The thicknesses of the significant paths reflect the magnitude of the standardized regression coefficients given alongside. R2-values inside boxes are conditional R2.
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